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Catalyzes the reduction of fatty acyl-CoA and -ACP (acyl carrier protein) substrates to fatty alcohols (PubMed:19062129, PubMed:27274541). Triggers the accumulation of C16 and C18 fatty alcohols; converts palmitoyl-acyl carrier protein to the corresponding C16:0 alcohol with NAD(P)H as electron donor, but seems inactive toward palmitoyl- or other acyl-coenzyme A (PubMed:21813653). Triggers also the formation of some C16:0 aldehydes (PubMed:27274541). Involved in the synthesis of the lipid component in sporopollenin (PubMed:19062129). Required for exine patterning of pollen grain by mediating the formation of pollen wall substances (PubMed:21813653, PubMed:21849515, PubMed:9351246).